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This is plausible given the high allelic diversity Other explanations such as historical introgression through hybridization and the small sample sizes of are also possible. Greater sampling effort south of the Yellow river will be necessary to differentiate between these explanations.

The observed allelic diversity in this study The greater average number of alleles per locus in this study is likely due to the different loci used and differences in microsatellite scoring between studies. Since this study had much more extensive sampling in this area it is not surprising that a greater diversity of alleles was found. The maintenance of and nuclear genomes in sympatry suggests that intrinsic barriers to gene flow exist, such as pre- or post-mating isolation mechanisms, preventing inter-breeding between these two species.

The majority of our sampling sites are from the overlap zone between the hypothesized ranges of the two species Figure 1. Furthermore, the two species are syntopic at sites 18, 19 and 21, and only one hybrid out of 54 individuals with an admixed nuclear genome was detected. If these were not two species one would expect most of the individuals examined to have admixed or hybrid genomes as a consequence of random mating between the two genotypes.

Nevertheless, the presence of individuals with admixed nuclear genomes suggests recent hybridization events between and and a lack of complete reproductive isolation. The genus is known for its complex social structure and mating behaviours, which may play an important role in mate choice and intrasexual communication. To date, the ecology and behaviour of these species remain poorly understood, and future work addressing the mating behaviour of these lizards will help elucidate potential isolation mechanisms between these species.

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